"Evolutionary dynamics" is the study of the fundamental mathematical principles that guide evolution. The Program for Evolutionary Dynamics (PED) at Harvard University was established in 2003 and is dedicated to research and teaching.

Martin Nowak, Professor of Mathematics and of Biology at Harvard University, is the director of this program. May Huang is the Chief Administrative Officer.

Our current research topics include: culturomics, eusociality, evolution of cooperation, cancer, viruses, evolutionary game/graph/set theory, happiness, networks, banks, prelife, spatial selection, and experimental games.


A brief statement about inclusive fitness and eusociality.

"Kin and Kind"




























David Rand et al. study how fairness prevails in uncertainty.
David Rand Kris Snibbe/Harvard Staff Photographer

Rapid acts of kindness: David Rand (center), Joshua Greene (left), and Martin Nowak (right).
Green, Rand, and Nowak Kris Snibbe/Harvard Staff Photographer

Alison Hill explains her forward thinking on HIV to Daniel Rosenbloom.
Alison Hill and Daniel Rosenbloom Jon Chase/Harvard Staff Photographer

Erez Lieberman Aiden and Jean-Baptiste Michel searching for n-grams.
Erez Lieberman Aiden and Jean-Baptiste Michel Kris Snibbe/Harvard Staff Photographer